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<html><head><meta http-equiv="Content-Type" content="text/html; charset=UTF-8"><title>SEC FORM 
            4</title><style type="text/css">
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      </style></head><body>SEC Form 4 
   <table width="100%" border="0" cellspacing="0" cellpadding="4"><tr><td width="20%" colspan="2" valign="top" align="center" class="FormName">FORM 4</td><td rowspan="2" width="60%" valign="middle" align="center"><span class="FormTitle">UNITED STATES SECURITIES AND EXCHANGE COMMISSION</span><br><span class="MedSmallFormText">Washington, D.C. 20549</span><br><br><span class="FormTitle">STATEMENT OF CHANGES IN BENEFICIAL OWNERSHIP</span><br><br><span class="MedSmallFormText">Filed pursuant to Section 16(a) of the Securities Exchange Act of 1934</span><br><span class="MedSmallFormText">or Section 30(h) of the Investment Company Act of 1940</span></td><td rowspan="2" width="20%" valign="top" align="center"><table width="100%" border="1" summary="OMB Approval Status Box"><tr><td class="FormTextC">OMB APPROVAL</td></tr><tr><td><table width="100%" border="0" summary="OMB Interior Box"><tr><td class="SmallFormText" colspan="3">OMB Number:</td><td class="SmallFormTextR">3235-0287</td></tr><tr><td class="SmallFormText" colspan="4">Estimated average burden</td></tr><tr><td class="SmallFormText" colspan="3">hours per response:</td><td class="SmallFormTextR">0.5</td></tr></table></td></tr></table></td></tr><tr valign="middle"><td><table width="100%" border="1" cellpadding="0" cellspacing="0"><tr><td>  </td></tr></table></td><td class="SmallFormText">Check this box if no longer subject to Section 16. Form 4 or Form 5 obligations may continue. 
         <i>See</i>

         Instruction 1(b).</td></tr><tr valign="middle"><td><table width="100%" border="1" cellpadding="0" cellspacing="0"><tr><td>  </td></tr></table></td><td class="SmallFormText">Check this box to indicate that a transaction was made pursuant to a contract, instruction or written plan for the purchase or sale of equity securities of the issuer that is intended to satisfy the affirmative defense conditions of Rule 10b5-1(c). See Instruction 10. </td></tr></table><table width="100%" border="1" cellspacing="0" cellpadding="4"><tr><td rowspan="4" width="35%" valign="top"><span class="MedSmallFormText">1. Name and Address of Reporting Person<sup>*</sup></span><table border="0" width="100%"><tr><td><a href="/cgi-bin/browse-edgar?action=getcompany&amp;CIK=0001726283">Johnson Natalia</a></td></tr></table><hr width="98%"><table border="0" width="100%"><tr><td width="33%" class="MedSmallFormText">(Last)</td><td width="33%" class="MedSmallFormText">(First)</td><td width="33%" class="MedSmallFormText">(Middle)</td></tr></table><table border="0" width="100%"><tr><td><span class="FormData">C/O PUBLIC STORAGE</span></td></tr><tr><td><span class="FormData">2811 INTERNET BOULEVARD</span></td></tr></table><hr width="98%"><span class="MedSmallFormText">(Street)</span><table border="0" width="100%"><tr><td width="33%"><span class="FormData">FRISCO</span></td><td width="33%"><span class="FormData">
				TEXAS
			</span></td><td width="33%"><span class="FormData">75034</span></td></tr></table><hr width="98%"><table border="0" width="100%"><tr><td width="33%" class="MedSmallFormText">(City)</td><td width="33%" class="MedSmallFormText">(State)</td><td width="33%" class="MedSmallFormText">(Zip)</td></tr></table><hr width="98%"><table border="0" width="100%"><tr><td width="33%" colspen="3"><span class="FormData">UNITED STATES</span></td></tr></table><hr width="98%"><span class="MedSmallFormText">(Country)</span></td><td width="35%" valign="top"><span class="MedSmallFormText">2. Issuer Name <b>and</b> Ticker or Trading Symbol
      </span><br><a href="/cgi-bin/browse-edgar?action=getcompany&amp;CIK=0001393311">Public Storage</a>
     [ <span class="FormData">PSA</span> ]
   </td><td rowspan="2" valign="top"><span class="MedSmallFormText">5. Relationship of Reporting Person(s) to Issuer
      </span><br><span class="MedSmallFormText">(Check all applicable)</span><table border="0" width="100%"><tr><td width="15%" align="center"></td><td width="35%" class="MedSmallFormText">Director</td><td width="15%" align="center"></td><td width="35%" class="MedSmallFormText">10% Owner</td></tr><tr><td align="center"><span class="FormData">X</span></td><td class="MedSmallFormText">Officer (give title below)</td><td align="center"></td><td class="MedSmallFormText">Other (specify below)</td></tr><tr><td width="15%" align="center"></td><td width="35%" align="left" style="color: blue">CD&amp;TO</td><td width="15%" align="center"></td><td width="35%" align="left" style="color: blue"></td></tr></table></td></tr><tr><td valign="top"><span class="MedSmallFormText">2a. Foreign Trading Symbol
		 <br></span></td></tr><tr><td valign="top"><span class="MedSmallFormText">3. Date of Earliest Transaction
         (Month/Day/Year)</span><br><span class="FormData">07/28/2026</span></td><td valign="top" rowspan="2"><span class="MedSmallFormText">6. Individual or Joint/Group Filing (Check Applicable Line)
      </span><table border="0" width="100%"><tr><td width="15%" align="center"><span class="FormData">X</span></td><td width="85%" class="MedSmallFormText">Form filed by One Reporting Person</td></tr><tr><td width="15%" align="center"></td><td width="85%" class="MedSmallFormText">Form filed by More than One Reporting Person</td></tr></table></td></tr><tr><td valign="top" rowspan="1"><span class="MedSmallFormText">4. If Amendment, Date of Original Filed
         (Month/Day/Year)</span><br><br></td></tr></table><table width="100%" border="1" cellspacing="0" cellpadding="4"><thead><tr><th width="100%" valign="top" colspan="11" align="center" class="FormTextC"><b>Table I - Non-Derivative Securities Acquired, Disposed of, or Beneficially Owned</b></th></tr><tr><th width="36%" valign="top" rowspan="2" align="left" class="MedSmallFormText">1. Title of Security (Instr. 
      3)
   </th><th width="6%" valign="top" rowspan="2" align="left" class="SmallFormText">2. Transaction Date
      (Month/Day/Year)</th><th width="5%" valign="top" rowspan="2" align="left" class="SmallFormText">2A. Deemed Execution Date, if any
      (Month/Day/Year)</th><th width="7%" valign="top" colspan="2" align="left" class="SmallFormText">3. Transaction Code (Instr. 
      8)
   </th><th width="19%" valign="top" colspan="3" align="left" class="SmallFormText">4. Securities Acquired (A) or Disposed Of (D) (Instr. 
      3, 4 and 5)
   </th><th width="11%" valign="top" rowspan="2" align="left" class="SmallFormText">5. 
      Amount of Securities Beneficially Owned Following Reported Transaction(s) (Instr. 
      3 and 4)
   </th><th width="9%" valign="top" rowspan="2" align="left" class="SmallFormText">6. Ownership Form: Direct (D) or Indirect (I) (Instr. 
      4)
   </th><th width="8%" valign="top" rowspan="2" align="left" class="SmallFormText">7. Nature of Indirect Beneficial Ownership (Instr. 
      4)
   </th></tr><tr><th width="4%" align="center" class="SmallFormText">Code</th><th width="3%" align="center" class="SmallFormText">V</th><th width="8%" align="center" class="SmallFormText">Amount</th><th width="5%" align="center" class="SmallFormText">(A) or (D)</th><th width="6%" align="center" class="SmallFormText">Price</th></tr></thead></table><table width="100%" border="1" cellspacing="0" cellpadding="4"><thead><tr><th width="100%" valign="top" colspan="16" align="center" class="FormTextC"><b>Table II - Derivative Securities Acquired, Disposed of, or Beneficially Owned</b><br><b>(e.g., puts, calls, warrants, options, convertible securities)</b></th></tr><tr><th width="13%" valign="top" rowspan="2" align="left" class="SmallFormText">1. Title of Derivative Security (Instr. 
      3)
   </th><th width="5%" valign="top" rowspan="2" align="left" class="SmallFormText">2. Conversion or Exercise Price of Derivative Security
   </th><th width="5%" valign="top" rowspan="2" align="left" class="SmallFormText">3. Transaction Date
      (Month/Day/Year)</th><th width="5%" valign="top" rowspan="2" align="left" class="SmallFormText">3A. Deemed Execution Date, if any
      (Month/Day/Year)</th><th width="9%" valign="top" colspan="2" align="left" class="SmallFormText">4. Transaction Code (Instr. 
      8)
   </th><th width="10%" valign="top" colspan="2" align="left" class="SmallFormText">5. 
      Number of Derivative Securities Acquired (A) or Disposed of (D) (Instr. 
      3, 4 and 5)
   </th><th width="9%" valign="top" colspan="2" align="left" class="SmallFormText">6. Date Exercisable and Expiration Date 
      (Month/Day/Year)</th><th width="17%" valign="top" colspan="2" align="left" class="SmallFormText">7. Title and Amount of Securities Underlying Derivative Security (Instr. 
      3 and 4)
   </th><th width="6%" valign="top" rowspan="2" align="left" class="SmallFormText">8. Price of Derivative Security (Instr. 
      5)
   </th><th width="6%" valign="top" rowspan="2" align="left" class="SmallFormText">9. 
      Number of derivative Securities Beneficially Owned Following Reported Transaction(s) (Instr. 
      4)
   </th><th width="6%" valign="top" rowspan="2" align="left" class="SmallFormText">10. Ownership Form: Direct (D) or Indirect (I) (Instr. 
      4)
   </th><th width="7%" valign="top" rowspan="2" align="left" class="SmallFormText">11. Nature of Indirect Beneficial Ownership (Instr. 
      4)
   </th></tr><tr><th width="4%" valign="bottom" align="center" class="SmallFormText">Code</th><th width="4%" valign="bottom" align="center" class="SmallFormText">V</th><th width="5%" valign="bottom" align="center" class="SmallFormText">(A)</th><th width="5%" valign="bottom" align="center" class="SmallFormText">(D)</th><th width="5%" valign="bottom" align="center" class="SmallFormText">Date Exercisable</th><th width="4%" valign="bottom" align="center" class="SmallFormText">Expiration Date</th><th width="10%" valign="bottom" align="center" class="SmallFormText">Title</th><th width="7%" valign="bottom" align="center" class="SmallFormText">Amount or Number of Shares</th></tr></thead><tbody><tr><td align="left"><span class="SmallFormData">AO LTIP Units</span></td><td align="center"><span class="SmallFormText">$</span><span class="SmallFormData">225.38</span><span class="FootnoteData"><sup>(1)</sup></span><span class="FootnoteData"><sup>(2)</sup></span></td><td align="center"><span class="SmallFormData">07/28/2026</span></td><td align="center"></td><td align="center"><span class="SmallFormData">M</span></td><td align="center"></td><td align="center"></td><td align="center"><span class="SmallFormData">10,327</span></td><td align="center"><span class="SmallFormData">08/05/2017</span><span class="FootnoteData"><sup>(1)</sup></span><span class="FootnoteData"><sup>(2)</sup></span></td><td align="center"><span class="SmallFormData">08/04/2026</span><span class="FootnoteData"><sup>(1)</sup></span><span class="FootnoteData"><sup>(2)</sup></span></td><td align="center"><span class="SmallFormData">Common Shares</span></td><td align="center"><span class="FormData">10,327</span></td><td align="center"><span class="FootnoteData"><sup>(1)</sup></span><span class="FootnoteData"><sup>(2)</sup></span></td><td align="center"><span class="SmallFormData">0</span></td><td align="center"><span class="SmallFormData">D</span></td><td align="left"></td></tr><tr><td align="left"><span class="SmallFormData">LTIP Units</span></td><td align="center"><span class="FootnoteData"><sup>(3)</sup></span></td><td align="center"><span class="SmallFormData">07/28/2026</span></td><td align="center"></td><td align="center"><span class="SmallFormData">M</span></td><td align="center"></td><td align="center"><span class="SmallFormData">3,284</span></td><td align="center"></td><td align="center"><span class="SmallFormData">
                
            </span><span class="FootnoteData"><sup>(3)</sup></span></td><td align="center"><span class="SmallFormData">
                
            </span><span class="FootnoteData"><sup>(3)</sup></span></td><td align="center"><span class="SmallFormData">Common Shares</span></td><td align="center"><span class="FormData">3,284</span></td><td align="center"><span class="SmallFormText">$</span><span class="SmallFormData">0</span></td><td align="center"><span class="SmallFormData">26,493</span><span class="FootnoteData"><sup>(4)</sup></span></td><td align="center"><span class="SmallFormData">D</span></td><td align="left"></td></tr></tbody></table><table border="0" width="100%"><tr><td class="MedSmallFormText"><b>Explanation of Responses:</b></td></tr><tr><td class="FootnoteData">1. On February 26, 2024, the reporting person exchanged an option to purchase 10,327 common shares of beneficial interest, par value $0.01 per share ("Common Shares"), of Public Storage (the "Company") for 10,327 limited partnership units in Public Storage OP, L.P. designated as AO LTIP Units ("AO LTIP Units"). AO LTIP Units are similar to "net exercise" stock option awards and are convertible, once vested, into a number of vested limited partnership units of Public Storage OP, L.P. designated as LTIP Units ("LTIP Units"), determined by the quotient of (i) the excess of the value of a Common Share as of the date of the conversion over $225.38, divided by (ii) the value of a Common Share as of the date of conversion. Vested LTIP Units into which AO LTIP Units have been converted are further convertible, [footnote continued]</td></tr><tr><td class="FootnoteData">2. [Continued from footnote] conditioned upon minimum allocations to the capital accounts of the LTIP Units for U.S. federal income tax purposes, into an equal number of limited partnership units in Public Storage OP, L.P. ("OP Units"). The resulting OP Units are redeemable by the holder for one Common Share per OP Unit or the cash value of a Common Share, at the Company's option.</td></tr><tr><td class="FootnoteData">3. Common Shares are issued upon the redemption of OP Units on a one for one basis. OP Units have no expiration date.</td></tr><tr><td class="FootnoteData">4. Includes 9,398 LTIP Units subject to time-based vesting.</td></tr><tr><td class="FormText"><b>Remarks:</b></td></tr><tr><td class="FootnoteData"></td></tr></table><table width="100%" border="0"><tr><td width="60%"></td><td width="20%"><u><span class="FormData">/s/ Nathaniel A. Vitan, Attorney-in-Fact</span></u></td><td width="20%"><u><span class="FormData">07/30/2026</span></u></td></tr><tr><td></td><td class="MedSmallFormText">** Signature of Reporting Person</td><td class="MedSmallFormText">Date</td></tr><tr><td colspan="3" class="MedSmallFormText">Reminder: Report on a separate line for each class of securities beneficially owned directly or indirectly.</td></tr><tr><td colspan="3" class="MedSmallFormText">* If the form is filed by more than one reporting person, 
                  <i>see</i>

                  Instruction 
                  4

                  (b)(v).</td></tr><tr><td colspan="3" class="MedSmallFormText">** Intentional misstatements or omissions of facts constitute Federal Criminal Violations 
                  <i>See</i>

                  18 U.S.C. 1001 and 15 U.S.C. 78ff(a).</td></tr><tr><td colspan="3" class="MedSmallFormText">Note: File three copies of this Form, one of which must be manually signed. If space is insufficient, 
                  <i>see</i>

                  Instruction 6 for procedure.</td></tr><tr><td colspan="3" class="MedSmallFormText"><b>Persons who respond to the collection of information contained in this form are not required to respond unless the form displays a currently valid OMB Number.</b></td></tr><tr><td colspan="3" class="MedSmallFormText"><b>* Form 4: SEC 1474 (03-26)</b></td></tr></table></body></html>
